LONDON (Reuters) - U.S. bank J.P. Morgan (>> JPMorgan Chase & Co.) pays managing directors at its investment bank in London an average of 461,000 pounds ($737,877), substantially more than pay across its rivals, according to a survey.

Emolument, a website that benchmarks salaries, said on Thursday the average salary and bonus for JPMorgan managing directors was more than 13 percent higher than the next best paying bank, Deutsche Bank (>> Deutsche Bank AG), which paid its managing directors an average of 402,000 pounds.

Bank of America Merrill Lynch (>> Bank of America Corp) paid an average of 376,000 pounds, according to Emolument, which said its estimates were based on responses from 640 bankers regarding their 2013/14 salaries and bonuses.

UBS (>> UBS AG), Citigroup (>> Citigroup Inc) and Morgan Stanley (>> Morgan Stanley) each paid managing directors an average of more than 350,000 pounds, the survey showed. Goldman Sachs (>> Goldman Sachs Group Inc), long considered one of the top paying banks, ranked ninth in the survey with average pay of 330,000, and Barclays ranked 13th.

(1 US dollar = 0.6248 British pound)

(Reporting by Steve Slater; Editing by Clara Ferreira Marques)